Jews

=== Genetic studies ===
{{main|Genetic studies on Jews}}
[[Y chromosome|Y DNA]] studies tend to imply a small number of founders in an old population whose members parted and followed different migration paths.<ref name="hammer2000">{{cite journal |last1=Hammer |first1=M. F. |last2=Redd |first2=A. J. |last3=Wood |first3=E. T. |last4=Bonner |first4=M. R. |last5=Jarjanazi |first5=H. |last6=Karafet |first6=T. |last7=Santachiara-Benerecetti |first7=S. |last8=Oppenheim |first8=A. |last9=Jobling |first9=M. A. |last10=Jenkins |first10=T. |last11=Ostrer |first11=H. |last12=Bonne-Tamir |first12=B. |title=Jewish and Middle Eastern non-Jewish populations share a common pool of Y-chromosome biallelic haplotypes |journal=Proceedings of the National Academy of Sciences |date=6 June 2000 |volume=97 |issue=12 |pages=6769–6774 |doi=10.1073/pnas.100115997 |pmid=10801975 |pmc=18733 |bibcode=2000PNAS...97.6769H |doi-access=free }}</ref> In most Jewish populations, these male line ancestors appear to have been mainly [[Middle East]]ern. For example, Ashkenazi Jews share more common paternal lineages with other Jewish and Middle Eastern groups than with non-Jewish populations in areas where Jews lived in [[Eastern Europe]], [[Germany]], and the French [[Rhine|Rhine Valley]]. This is consistent with Jewish traditions in placing most Jewish paternal origins in the region of the Middle East.<ref name="Nebel 2001">{{cite journal |last1=Nebel |first1=Almut |last2=Filon |first2=Dvora |last3=Brinkmann |first3=Bernd |last4=Majumder |first4=Partha P. |last5=Faerman |first5=Marina |last6=Oppenheim |first6=Ariella |title=The Y Chromosome Pool of Jews as Part of the Genetic Landscape of the Middle East |journal=The American Journal of Human Genetics |date=November 2001 |volume=69 |issue=5 |pages=1095–1112 |doi=10.1086/324070 |pmid=11573163 |pmc=1274378 }}</ref><ref>{{cite book |last1=Frudakis |first1=Tony |chapter=Ashkezani Jews |page=383 |chapter-url=https://books.google.com/books?id=9vXeydpj7VkC&pg=PA383 |title=Molecular Photofitting: Predicting Ancestry and Phenotype Using DNA |date=19 July 2010 |publisher=Elsevier |isbn=978-0-08-055137-1 }}</ref>

Conversely, the maternal lineages of Jewish populations, studied by looking at [[mitochondrial DNA]], are generally more heterogeneous.<ref name="Behar2008b">{{cite journal |last1=Behar |first1=Doron M. |last2=Metspalu |first2=Ene |last3=Kivisild |first3=Toomas |last4=Rosset |first4=Saharon |last5=Tzur |first5=Shay |last6=Hadid |first6=Yarin |last7=Yudkovsky |first7=Guennady |last8=Rosengarten |first8=Dror |last9=Pereira |first9=Luisa |last10=Amorim |first10=Antonio |last11=Kutuev |first11=Ildus |last12=Gurwitz |first12=David |last13=Bonne-Tamir |first13=Batsheva |last14=Villems |first14=Richard |last15=Skorecki |first15=Karl |title=Counting the Founders: The Matrilineal Genetic Ancestry of the Jewish Diaspora |journal=PLOS ONE |date=30 April 2008 |volume=3 |issue=4 |pages=e2062 |doi=10.1371/journal.pone.0002062 |pmid=18446216 |pmc=2323359 |bibcode=2008PLoSO...3.2062B |doi-access=free }}</ref> Scholars such as [[Harry Ostrer]] and [[Raphael Falk (academic)|Raphael Falk]] believe this indicates that many Jewish males found new mates from European and other communities in the places where they migrated in the diaspora after fleeing ancient Israel.<ref name="Lewontin">{{Cite journal |last=Lewontin |first=Richard |author-link=Richard Lewontin |date=6 December 2012 |title=Is There a Jewish Gene? |url=http://www.nybooks.com/articles/2012/12/06/is-there-a-jewish-gene/ |journal=New York Review of Books|volume=59 |issue=19 }}</ref> In contrast, Behar has found evidence that about 40&nbsp;percent of Ashkenazi Jews originate maternally from just four female founders, who were of Middle Eastern origin. The populations of Sephardi and Mizrahi Jewish communities "showed no evidence for a narrow founder effect."<ref name="Behar2008b" /> Subsequent studies carried out by Feder et al. confirmed the large portion of non-local maternal origin among Ashkenazi Jews. Reflecting on their findings related to the maternal origin of Ashkenazi Jews, the authors conclude "Clearly, the differences between Jews and non-Jews are far larger than those observed among the Jewish communities. Hence, differences between the Jewish communities can be overlooked when non-Jews are included in the comparisons."<ref name="Abraham 2010"/><ref>{{cite journal |last1=Feder |first1=Jeanette |last2=Ovadia |first2=Ofer |last3=Glaser |first3=Benjamin |last4=Mishmar |first4=Dan |title=Ashkenazi Jewish mtDNA haplogroup distribution varies among distinct subpopulations: lessons of population substructure in a closed group |journal=European Journal of Human Genetics |date=April 2007 |volume=15 |issue=4 |pages=498–500 |doi=10.1038/sj.ejhg.5201764 |pmid=17245410 |doi-access=free}}</ref><ref name="ReferenceA">{{cite journal |last1=Ostrer |first1=Harry |last2=Skorecki |first2=Karl |title=The population genetics of the Jewish people |journal=Human Genetics |date=February 2013 |volume=132 |issue=2 |pages=119–127 |doi=10.1007/s00439-012-1235-6 |pmid=23052947 |pmc=3543766 }}</ref> A study showed that 7% of Ashkenazi Jews have the haplogroup G2c, which is mainly found in [[Pashtuns]] and on lower scales all major Jewish groups, Palestinians, Syrians, and Lebanese.<ref>{{cite web | title=Sign In | website=Family Tree DNA | url=https://www.familytreedna.com/sign-in?ReturnUrl=%2Fpdf%2FBehar_contrasting.pdf | access-date=1 April 2018}}</ref><ref>{{cite journal |last1=Hammer |first1=Michael F. |last2=Behar |first2=Doron M. |last3=Karafet |first3=Tatiana M. |last4=Mendez |first4=Fernando L. |last5=Hallmark |first5=Brian |last6=Erez |first6=Tamar |last7=Zhivotovsky |first7=Lev A. |last8=Rosset |first8=Saharon |last9=Skorecki |first9=Karl |title=Extended Y chromosome haplotypes resolve multiple and unique lineages of the Jewish priesthood |journal=Human Genetics |date=8 August 2009 |volume=126 |issue=5 |pages=707–17 |doi=10.1007/s00439-009-0727-5 |pmid=19669163 |pmc=2771134 }}</ref>

Studies of [[Autosome|autosomal DNA]], which look at the entire DNA mixture, have become increasingly important as the technology develops. They show that Jewish populations have tended to form relatively closely related groups in independent communities, with most in a community sharing significant ancestry in common.<ref>{{cite journal |last1=Katsnelson |first1=Alla |title=Jews worldwide share genetic ties |journal=Nature |date=3 June 2010 |pages=news.2010.277 |doi=10.1038/news.2010.277 }}</ref> For Jewish populations of the diaspora, the genetic composition of [[Ashkenazi Jews|Ashkenazi]], [[Sephardic Jews|Sephardic]], and [[Mizrahi Jews|Mizrahi]] Jewish populations show a predominant amount of shared Middle Eastern ancestry. According to Behar, the most parsimonious explanation for this shared Middle Eastern ancestry is that it is "consistent with the historical formulation of the Jewish people as descending from ancient [[Hebrews|Hebrew]] and [[Israelites|Israelite]] residents of the [[Levant]]" and "the dispersion of the people of ancient Israel throughout the [[Old World]]".<ref name="discovermagazine">{{cite journal |last1=Behar |first1=Doron M. |last2=Yunusbayev |first2=Bayazit |last3=Metspalu |first3=Mait |last4=Metspalu |first4=Ene |last5=Rosset |first5=Saharon |last6=Parik |first6=Jüri |last7=Rootsi |first7=Siiri |last8=Chaubey |first8=Gyaneshwer |last9=Kutuev |first9=Ildus |last10=Yudkovsky |first10=Guennady |last11=Khusnutdinova |first11=Elza K. |last12=Balanovsky |first12=Oleg |last13=Semino |first13=Ornella |last14=Pereira |first14=Luisa |last15=Comas |first15=David |last16=Gurwitz |first16=David |last17=Bonne-Tamir |first17=Batsheva |last18=Parfitt |first18=Tudor |last19=Hammer |first19=Michael F. |last20=Skorecki |first20=Karl |last21=Villems |first21=Richard |title=The genome-wide structure of the Jewish people |journal=Nature |date=July 2010 |volume=466 |issue=7303 |pages=238–242 |doi=10.1038/nature09103 |pmid=20531471 |s2cid=4307824 |bibcode=2010Natur.466..238B }}</ref> [[North Africa]]n, [[Italian Peninsula|Italian]] and others of [[Iberian Peninsula|Iberian]] origin show variable frequencies of admixture with non-Jewish historical host populations among the maternal lines. In the case of Ashkenazi and Sephardi Jews (in particular [[Moroccan Jews]]), who are closely related, the source of non-Jewish admixture is mainly [[Southern Europe]]an, while Mizrahi Jews show evidence of admixture with other Middle Eastern populations. Behar ''et al.'' have remarked on a close relationship between Ashkenazi Jews and modern [[Italians]].<ref name="discovermagazine"/><ref name=zooss>{{cite journal |last1=Zoossmann-Diskin |first1=Avshalom |title=The origin of Eastern European Jews revealed by autosomal, sex chromosomal and mtDNA polymorphisms |journal=Biology Direct |date=2010 |volume=5 |issue=1 |pages=57 |doi=10.1186/1745-6150-5-57 |pmid=20925954 |pmc=2964539 |bibcode=2010Sci...328.1342B |doi-access=free }}</ref> A 2001 study found that Jews were more closely related to groups of the Fertile Crescent (Kurds, Turks, and Armenians) than to their Arab neighbors, whose genetic signature was found in geographic patterns reflective of Islamic conquests.<ref name="Nebel 2001"/><ref>{{cite journal |last1=Haber |first1=Marc |last2=Gauguier |first2=Dominique |last3=Youhanna |first3=Sonia |last4=Patterson |first4=Nick |last5=Moorjani |first5=Priya |last6=Botigué |first6=Laura R. |last7=Platt |first7=Daniel E. |last8=Matisoo-Smith |first8=Elizabeth |last9=Soria-Hernanz |first9=David F. |last10=Wells |first10=R. Spencer |last11=Bertranpetit |first11=Jaume |last12=Tyler-Smith |first12=Chris |last13=Comas |first13=David |last14=Zalloua |first14=Pierre A. |title=Genome-Wide Diversity in the Levant Reveals Recent Structuring by Culture |journal=PLOS Genetics |date=28 February 2013 |volume=9 |issue=2 |pages=e1003316 |doi=10.1371/journal.pgen.1003316 |pmid=23468648 |pmc=3585000 |doi-access=free }}</ref>

The studies also show that [[Sephardic Bnei Anusim]] (descendants of the "[[anusim]]" who were [[Forced conversion|forced to convert]] to [[Catholicism]]), which comprise up to 19.8&nbsp;percent of the population of today's [[Iberia]] ([[Spain]] and [[Portugal]]) and at least 10&nbsp;percent of the population of [[Ibero-America]] ([[Hispanic America]] and [[Brazil]]), have Sephardic Jewish ancestry within the last few centuries. The [[Bene Israel]] and [[Cochin Jews]] of [[India]], [[Beta Israel]] of [[Ethiopia]], and a portion of the [[Lemba people]] of [[Southern Africa]], despite more closely resembling the local populations of their native countries, have also been thought to have some more remote ancient Jewish ancestry.<ref>{{cite web|url=http://forward.com/articles/155742/jews-are-a-race-genes-reveal/?p=all |title=Jews Are a 'Race,' Genes Reveal |date=4 May 2012 |publisher=Forward.com |access-date=12 April 2013}}</ref><ref name=discovermagazine/><ref name="in.reuters.com">{{cite news |last1=Begley |first1=Sharon |title=Genetic study offers clues to history of North Africa's Jews |url=https://in.reuters.com/article/us-science-genetics-jews/genetic-study-offers-clues-to-history-of-north-africas-jews-idINBRE8751EI20120806 |work=Reuters |date=6 August 2012 |access-date=12 October 2020 |archive-date=18 November 2015 |archive-url=https://web.archive.org/web/20151118100801/http://in.reuters.com/article/2012/08/06/us-science-genetics-jews-idINBRE8751EI20120806 |url-status=dead }}</ref><ref name="ReferenceA"/> Views on the Lemba have changed and genetic [[Y-DNA]] analyses in the 2000s have established a partially Middle-Eastern origin for a portion of the male Lemba population but have been unable to narrow this down further.<ref name="SpurdleJenkins">{{Citation | title = The origins of the Lemba "Black Jews" of southern Africa: evidence from p12F2 and other Y-chromosome markers. | pmid = 8900243 | pmc=1914832 | volume=59 | issue = 5 | date=November 1996 | journal=Am. J. Hum. Genet. | pages=1126–33 | last1 = Spurdle | first1 = AB | last2 = Jenkins | first2 = T}}</ref><ref name="Soodyall">{{cite book|author1=Himla Soodyall|author2=Jennifer G. R Kromberg|editor1-last=Kumar|editor1-first=Dhavendra|editor2-last=Chadwick|editor2-first=Ruth|title=Genomics and Society: Ethical, Legal, Cultural and Socioeconomic Implications|publisher=Academic Press/Elsevier|isbn=978-0-12-420195-8|page=316|chapter-url=https://books.google.com/books?id=E9icBAAAQBAJ&q=Cohen+Modal+Haplotype+Lemba&pg=PA309|chapter=Human Genetics and Genomics and Sociocultural Beliefs and Practices in South Africa|date=29 October 2015}}</ref>